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The head and body length of three-toed sloths averages 58 cm 23 inches , and the tail is short, round, and movable. The forelimbs are 50 percent longer than the hind limbs; all four feet have three long, curved sharp claws.
The outer layer of shaggy long hair is pale brown to gray and covers a short, dense coat of black-and-white underfur.
The outer hairs have many cracks, perhaps caused by the algae living there. The algae give the animals a greenish tinge, especially during the rainy season.
Sexes look alike in the maned sloth, but in the other species males have a large patch speculum in the middle of the back that lacks overhair, thus revealing the black dorsal stripe and bordering white underfur, which is sometimes stained yellow to orange.
The maned sloth gets its name from the long black hair on the back of its head and neck. Three-toed sloths, although mainly nocturnal, may be active day or night but spend only about 10 percent of their time moving at all.
They sleep either perched in the fork of a tree or hanging from a branch, with all four feet bunched together and the head tucked in on the chest.
In this posture the sloth resembles a clump of dead leaves, so inconspicuous that it was once thought these animals ate only the leaves of cecropia trees because in other trees it went undetected.
Research has since shown that they eat the foliage of a wide variety of other trees and vines. Locating food by touch and smell, the sloth feeds by hooking a branch with its claws and pulling it to its mouth.
Normally, three-toed sloths are silent and docile , but if disturbed they can strike out furiously with the sharp foreclaws. Reproduction is seasonal in the brown- and pale-throated species; the maned sloth may breed throughout the year.
Sloths are a group of arboreal Neotropical xenathran mammals, constituting the suborder Folivora. Noted for their slowness of movement, they spend most of their lives hanging upside down in the trees of the tropical rainforests of South America and Central America.
They are considered to be most closely related to anteaters , together making up the xenarthran order Pilosa. There are six extant sloth species in two genera — Bradypus three—toed sloths and Choloepus two—toed sloths.
Despite this traditional naming, all sloths actually have three toes on each rear limb, although two-toed sloths have only two digits on each forelimb.
Besides the extant species, many species of ground sloths ranging up to the size of elephants like Megatherium inhabited both North and South America during the Pleistocene Epoch.
However, they became extinct during the Quaternary extinction event around 12, years ago, together with most large bodied animals in the New World.
The extinction correlates in time with the arrival of humans, but climate change has also been suggested to have contributed.
Members of an endemic radiation of Caribbean sloths formerly lived in the Greater Antilles. They included both ground and arboreal forms which became extinct after humans settled the archipelago in the mid-Holocene, around 6, years ago.
Sloths are so named because of their very low metabolism and deliberate movements, sloth being related to the word slow. This supports their low-energy diet of leaves and avoids detection by predatory hawks and cats that hunt by sight.
The algae also nourishes sloth moths , some species of which exist solely on sloths. Sloths belong to the superorder Xenarthra , a group of placental mammals believed to have evolved in the continent of South America around 60 million years ago.
The earliest xenarthrans were arboreal herbivores with sturdy vertebral columns , fused pelvises , stubby teeth, and small brains. Sloths are in the taxonomic suborder Folivora  of the order Pilosa.
These names are from the Latin 'leaf eater' and 'hairy', respectively. Pilosa is one of the smallest of the orders of the mammal class; its only other suborder contains the anteaters.
The common ancestor of the two existing sloth genera dates to about 28 million years ago,  with similarities between the two- and three- toed sloths an example of convergent evolution to an arboreal lifestyle, "one of the most striking examples of convergent evolution known among mammals".
Ancient sloths were mostly terrestrial, and some reached sizes that rival those of elephants, as was the case for Megatherium. Sloths arose in South America during its long period of isolation and eventually spread to a number of the Caribbean islands as well as North America.
It is thought that swimming led to oceanic dispersal of pilosans to the Greater Antilles by the Oligocene , and that the megalonychid Pliometanastes and the mylodontid Thinobadistes were able to colonise North America about 9 million years ago, well before the formation of the Isthmus of Panama.
The latter development, about 3 million years ago, allowed megatheriids and nothrotheriids to also invade North America as part of the Great American Interchange.
Additionally, the nothrotheriid Thalassocnus of the west coast of South America became adapted to a semiaquatic and, eventually, perhaps fully aquatic marine lifestyle.
Initially they just stood in the water, but over a span of 4 million years they eventually evolved into swimming creatures, becoming specialist bottom feeders of seagrasses , similar to extant marine sirenians.
Both types of extant tree sloth tend to occupy the same forests; in most areas, a particular species of the somewhat smaller and generally slower-moving three-toed sloth Bradypus and a single species of the two-toed type will jointly predominate.
Based on morphological comparisons, it was thought the two-toed sloths nested phylogenetically within one of the divisions of the extinct Greater Antilles sloths.
Recently obtained molecular data from collagen  and mitochondrial DNA sequences  fall in line with the diphyly convergent evolution hypothesis, but have overturned some of the other conclusions obtained from morphology.
These investigations consistently place two-toed sloths close to mylodontids and three-toed sloths within Megatherioidea, close to Megalonyx , megatheriids and nothrotheriids.
They make the previously recognized family Megalonychidae polyphyletic , with both two-toed sloths and Greater Antilles sloths being moved away from Megalonyx.
Greater Antilles sloths are now placed in a separate, basal branch of the sloth evolutionary tree. The following sloth family phylogenetic tree is based on collagen and mitochondrial DNA sequence data see Fig.
Bradypodidae three-toed sloths. Choloepodidae two-toed sloths. The marine sloths of South America's Pacific coast became extinct at the end of the Pliocene following the closing of the Central American Seaway ; this caused a cooling trend in the coastal waters which killed off much of the area's seagrass and which would have also made thermoregulation difficult for the sloths, with their slow metabolism.
Ground sloths disappeared from both North and South America shortly after the appearance of humans about 11, years ago.
Evidence suggests human hunting contributed to the extinction of the American megafauna. Ground sloth remains found in both North and South America indicate that they were killed, cooked, and eaten by humans.
Megalocnus and some other Caribbean sloths survived until about 5, years ago, long after ground sloths had died out on the mainland, but then went extinct when humans finally colonized the Greater Antilles.
Two-toed sloths are slightly larger. Sloths are unusual among mammals in not having seven cervical vertebrae.
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